158 p. ill. Proefschrift Vrije Universiteit (VU) Amsterdam, Department of Theoretical Biology and the Research Institute for Plant Protection (IPO-DLO) Financially supported by the Ministry of Transport, Public Works and Water Management = Ministerie van Verkeer en Waterstaat, Rijkswaterstaat, National Institute for Coastal and Marine Management = Rijksinstituut voor Kust en Zee (RWS, RIKZ), within the framework of the programme System Analysis Western Scheldt Estuary (SAWES) and by the Ministry of Agriculture, Fisheries and Nature Management = Ministerie van Landbouw, Natuurbeheer en Visserij (LNV) With ref.